• Root
• Nature
• Animals
• Arthropods
• Insects
• Beetles
• Lucaninae
• Lucanidae
• Lucanini
• Dorcus

	Parent	Category	Figures
1	titanus	sika	5
2	titanus	sakishimanus	2
3	titanus	castanicolor	2
4	titanus	elegans	2
5	titanus	pilifer	1